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OREODONTS  OF  THE  TICK  CANYON  FORMATION,
                                      SOUTHERN  CALIFORNIA

                                     BY  DAVID  P.  WHISTLER

                   In  1940,  Richard  H.  Jahns  reviewed  the  stratigraphy  of
             the  nonmarine  Mint  Canyon  Formation  in  the  eastern  part  of
             the  Ventura  Basin  and  separated  from  it  a  new  formation  and
             fauna.   Previous  to  this  work,  a  controversy  existed  as  to
             the  age  of  the  Mint  Canyon  Formation,  for  it  contained
             vertebrates  considered  indicative  of  both  the  Miocene  and
             Pliocene  (Kew,  1924,  Maxson,  1930,  and  Stirton,  1933).  As  a
             partial  solut,ion  to  this  controversy,  Jahns  demonstrated  the
             presence  of  an  erosional  unconformity  low  in  the  nonmarine
             sequence  which  indicated  a  shift  in  source  area.  He
             redefined  the  beds  below  the  unconformity  as  the  Tick  Canyon
             Formation  (Jahns,  1940,  pp.  163-66).
                   Additional  fossils  discovered  in  the  Tick  Canyon
             Formation  since  Maxson's  work,  and  certain  of  the  forms
             described  by  Maxson,  comprise  the  Tick  Canyon  fauna.      Only
             two  specimens  described  by  Maxson  are  from  the  Tick  Canyon
             Formation,  UCMP  30046,  the  type  of  Miolabis  californicus
             and  UCMP  23852,  a  dentary  fragment  of  a  Parahippus.  Neither
             of  these  permitted  a  definitive  age  determination.       The
             additional  fauna  described  by  Jahns  indicates  an  Arikareean
             mammalian  age  (early  Miocene),  and  there  is  a  noteworthy
             temporal  hiatus  between  the  Tick  Canyon  fauna  and  the  over-
             lying  Mint  Canyon  fauna.    In  addition,  faunas  comprising
             three  mammalian  ages,  late  Barstovian,  and  earlier  and  late
             Clarendonian,  are  now  recognized  from  the  Mint  Canyon
             Formation,  but  this  is  not  the  principal  concern  of  this
             paper.
                   Several  reviews  of  individual  taxa  from  the  Tick  Canyon
             fauna  describe  either  new  genera  or  species  (Dawson,  1958,
             Reeder,  1960).  All  the  described  taxa  were  originally
             restricted  to  the  Tick  Canyon  Formation.      Unpublished  records
             of  these  taxa  now  occur  in  other  California  localities.
                   The  composite  Tick  Canyon  fauna  as  presently  known  is
             as  follows:    a  heteromyid  rodent,  Trogomys  rupinimenthae
             Reeder,  1960,  a  rabbit,  Archaeolagus  acaricolus  Dawson,
             1958,  two  oreodonts,  Merychyus  (Merychyus)  calaminthus  Jahns,
             1940,  and  Merychyus  (Merychyus)  jahnsi  (this  paper),  a  camel,


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